Bursera
longipes
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This
beautiful tree is found in the central Balsas Depression in central
southern
Mexico. It grows in very dry
forest with many other Bursera species and many columnar cacti.
The middle photo from atop a B. longipes while
taking
measurements of the branches, and shows the columnar cactus Neobuxbaumia
and two other Bursera species. The photo right of
center is of a tropical dry forest in which B. longipes is one of the most
common trees. You can see the red to copper bark and bluish foliage
here and there across the hillside. The largest individual we have
seen, at lower right, is in the parking lot at the Xochicalco
archeological site in Morelos state. |
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Bursera
grandifolia |
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With large,
arcing
trunks that branch near the base, this species looks like an enormous
rounded
shrub. Bursera grandifolia does indeed have very large
leaflets,
which tend to be very fuzzy. The species is found in tropical dry
forests
of central Mexico. The greenish gray inner bark contrasts strongly with
the
reddish outermost bark, making this species a striking member of the
dry
forest flora. |
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The
flowers of Bursera species
are usually produced in male and female versions, though there are
often exceptions. This photo shows the female flowers of B. grandifolia, which have three
petals.
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The
male flowers, in contrast, have five petals.
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Bursera
instabilis |
This
strange
species from western Mexico begins life like a normal tree, with a
conventional
trunk and branches that all support their own weight (left). With age,
the
branches begin to grow back on themselves, growing around and around
within
the crown of the tree. These branches rest on earlier-formed,
self-supporting
branches, much in the way that a liana would. One such branch removed
from
the tree is shown at right. It was 8 cm in diameter and 4 meters
long! |
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Bursera
arborea |
This
species
is remarkable in the simaruba clade for being the species with
the
thinnest twigs. Whereas all of the other species have very thick twigs,
the
young shoots of B. arborea are remarkably slender. The species
is
found in tropical dry forest on the Pacific coast of Mexico. These
photos
were taken in Jalisco state.
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Bursera
cinerea |
This
species
grows on the fringes of lowland dry tropical areas, at the transition
to
moister highland vegetation
in Oaxaca and Veracruz.
These magnificent trees are particularly striking for the deep red
color
of their trunks, which are an amazing sight sprawling in deep, shady
canyons,
as at left, or ascending in open forest, as at right. |
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Bursera
roseana |
Another
large, B. simaruba-like tree, this species was formerly known
as Terebinthus acuminatus. Like some other members of the simaruba
clade, this species grows in moist canyons in the transition
zone
between highland pine-oak forest and lowland tropical subdeciduous
forest. It is very hard to tell apart from B. simaruba and B. attenuata. |
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Bursera
attenuata |
This
species
is similar in many respects to B. simaruba, differing in some
leaf
characteristics. It grows in fairly tall, moist tropical subdeciduous
forests
of the Pacific coast of Mexico. In some places, it occurs very close to
sharp
transition zones, with pines and oaks growing just above a zone of
broadleaf
forest. |
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Bursera
simaruba |
This is
the
most widespread species of Bursera, occurring from southern
Florida
and the Caribbean, along both coasts of Mexico to South America. The
species
is highly variable and is found in everything from tropical dry forest
to
tall rainforest. The images in the upper row are from rainforest on the
Gulf coast of
Mexico. |
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Bursera
simaruba
as a landscape plant in the Comalcalco archaeological site in Tabasco.
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Bursera simaruba pruned and shaped
by salt laden sea breezes just above the beach in Veracruz. |
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Bursera simaruba is often used as
a living fence. Thick sections of branch readily root and make perfect
fence posts that don't rot in tropical climates as dead wooden posts
would. This fence is very close to the Gulf in Veracruz.
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Bursera
simaruba is
also found on the Pacific slope, from Sonora to South America. This
photo is from a shady arroyo in the Chamela research station on the
coast of Jalisco. The photos in the row below are also from the coast
of Jalisco and Nayarit, where B.
simaruba
often grows in tall shady forests with Orbigyna palms endemic to the
central Mexican Pacific.
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In Mexico, the
Pacific and Atlantic populations of Bursera
simaruba are widely separated. Where the land narrows in Central
America, these plants can be found on both coasts very close to one
another in very different forest types, from Caribbean slope rainforest
to Pacific dry forest, as in these images are from Costa Rica. At left
are trees from the Caribbean, at Cahuita, and at the Palo Verde
research station on the Pacific in the two remaining images.
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Bursera simaruba is found on all
the islands of the Caribbean, from the smallest to the largest. Here,
we show it growing with the cycad Microcycas
calocoma in dense woods on limestone.
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Bursera laurihuertae |
Our research shows that this enigmatic
species is a member of the simaruba complex. It is endemic to the low,
dense tropical dry forests of the southern Isthmus of Tehuantepec and
is very distinctive for its rounded, leathery leaves that have just
three or even only one leaflet. |
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Bursera ovalifolia |
This species was only recently recognized as
an entitiy distinct from Bursera
simaruba. It grows in warm, rather moist tropical deciduous
forests on the Pacific coast of Mexico. It is sometimes hard to tell
from B. simaruba but has
longer petiolules and much smaller fruits. The three photos on the left
in the row below are from Guerrero; the one on the right is from the
central dry depression of Chiapas.
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We visited the type locality of
Bursera longicuspis in
Chiapas, to find that the plants there are individuals of B. ovalifolia growing at the upper
elevational extent of the species, in the transition to oak forest. The
plants seemed shorter and squatter than those at lower elevations.
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Bursera
species
usually flower and start to leaf out just as the dry season is ending.
This Bursera
ovalifolia
in the forest of the Chamela resarch station in Jalisco is crowned with
leaves in the first week of rain of the season. The rest of the trees
in the forest are not far behind.
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Bursera krusei |
This species is very distinctive for its
leaves with three large hairy leaflets. It grows in dry tropical forest
in Guerrero and Oaxaca.
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Bursera
standleyana
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This remarkable species is a
hemiepiphyte. That is, it grows high in other trees, but sends long,
flexible, liana-like roots down to the forest floor. It is known only
from the moist forest of the central and southern Pacific slopes of
Costa Rica. In the photo with arrows, the upper arrow indicates one of
the arching branches of B.
standleyana and the lower one highlights the supporting tree.
The leaves of B. standleyana can
have five, three, or even just one leaflet.
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Bursera
shaferi
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The group of Bursera species
that are endemic to the islands of the Caribbean appears to be part of
the simaruba clade. Bursera shaferi is
a little-known small tree that only grows on limestone mountains in
Pinar del Río. |